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Your specific board has bandwidth sharing between PCI E slots
PCIE 16x 1 slot Halves its bandwidth when PCIE 8x 1 slot is occupied and PCIE 16x 2 slot Halves its bandwidth when PCIE 8x 2 slot is occupied. So that means to prevent your 16x slots from losing half their bandwidth your 8x slots have to remain empty, no matter what card you have in the 8x slot.
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